Habitats: temperate broadleaf/mixed forest (25), tropical/subtropical dry broadleaf forest (3)
Found in 31 samples
China: Changbai Mountain Reserve, Baihua, Lühe Town (Early Oligocene)
Denmark: Bolderslev Skov
Hungary: Wind Brickyard (Late Oligocene)
India: Mawnai
Japan: Shiretoko National Park, University of Tokyo Hokkaido Forest
Mexico: Northeastern Coastal Plain of Coahuila (Quercus fusiformis forest), Northeastern Coastal Plain of Coahuila (Carya illinoinensis forest)
United States: Missouri River Floodplain, Abner's Hollow, Sunrise Towhead Island, Stewart State Forest (H), Stewart State Forest (NH), North Woods (oak-hickory community), North Woods (transition community), North Woods (maple-linden community), Cedar Creek Natural History Area, Waubun, Hill Forest State Natural Area, Rich Harvest Links, Beadles Barrens Nature Preserve, Bonayer Forest, Sipsey Wilderness, Pogue Creek Natural Area, Wharton Brook, Botany Glen (Plot 1), Botany Glen (Plot 2), Botany Glen (Plot 3)
Vietnam: Yen Bai (Late Miocene)
See also Ulmus alata, Ulmus americana, Ulmus crassifolia, Ulmus davidiana, Ulmus fischeri, Ulmus fulva, Ulmus glabra, Ulmus laciniata, Ulmus lanceifolia, Ulmus macrocarpa, Ulmus propinqua, Ulmus pseudopyramidalis, Ulmus pyramidalis, Ulmus rubra, Ulmus serotina, Ulmus sp., Ulmus spp.