Coudoulous I (Faunal Unit II)
Basic information
Sample name: Coudoulous I (Faunal Unit II)

Reference: P. Fernandez, M. Jeannet, J. Jaubert, and J.-P. Brugal. 2021. Paleoecological first results on the Middle Pleistocene sequence of Coudoulous I (Quercy, Lot, France). Alpine and Mediterranean Quaternary 34(1):89-107 [ER 3857]
Geography
Country: France

Coordinate: 44° 28' 20" N, 2° 40' 50" E
Latlng basis: stated in text

Time interval: Middle Pleistocene

Max Ma: 0.229

Min Ma: 0.1613

Age basis: other

Geography comments: "in the Quercy area... at the junction of the river LOt and its tributary, the Célé"
the entire faunal sequence is bracketed by an ESR/U-series date of 161.3 +/- 14 kyr at the top and two U/Th dates of 229 + 78 - 42 and 202 + 78 - 39 kyr at the base

Environment
Lithology: siliciclastic (mixed)

Taphonomic context: cave, human accumulation, pitfall trap

Archaeology: stone tools

Habitat comments: a pitfall trap with fossils from "clays, sands, angular limestone rocks"
there is a "very rich archaeological level" immediately above the fossiliferous sequence that dates to the "Early Middle Paleolithic" and there are "scarce and dispersed artefacts of the same age" in the fossiliferous levels
the site "was used for marginal scavenging by Lower Paleolithic hominid groups and as a kill-butchery site during the EMP", so it is at least partially a human accumulation
this sample is from stratigraphic unit 5
the material may mostly be from the sandstone comprising unit 5b, but may also be from the thinner apparent clay in unit 5a; both include breccia

Methods
Life forms: bats, carnivores, rodents, ungulates, other small mammals, lizards, snakes, frogs

Sampling methods: quarry, screenwash

Sample size: 2024 specimens

Years: 1994 - 2003

Net or trap nights: 0

Basal area status: not applicable

Sampling comments: there was a "rescue excavation" in 1978 - 1980, but the main excavation was between 1994 and 2003
the high abundance of micromammals is taken to mean that screening was used

Metadata
Sample no: 4223

Contributor no: John Alroy

Enterer: John Alroy

Created: 2023-04-13 17:27:15

Modified: 2023-04-13 07:27:15

Abundance distribution
Each square represents a species. Square sizes are proportional to counts. Values are logged.
Statistics
53 species
18 singletons
total count 2024
geometric series index: 99.1
Fisher's α: 9.965
geometric series k: 0.875
Hurlbert's PIE: 0.7157
Shannon's H: 2.0626
Good's u: 0.9911
Register
Microtus arvalis 1037
Microtus agrestis 168
Stenocranius gregalis 98
"Microtus gregalis"
Alexandromys oeconomus (tundra vole)20
"Microtus oeconomus"
Chionomys nivalis 1
Microtus brecciensis 88
Microtus multiplex 68
Microtus gerbii 41
"Microtus pyrenaicus"
Microtus subterraneus 42
Clethrionomys glareolus (bank vole)7
Pliomys episcopalis 1
Pliomys lenki 4
Arvicola cantiana (vole)7
Allocricetus bursae 32
Apodemus sylvaticus (wood mouse)163
Apodemus flavicollis (yellow-necked mouse)10
Apodemus uralensis (Ural field mouse)36
Eliomys quercinus 44
Sciurus vulgaris (Eurasian red squirrel)1311.0 g
Talpa europaea (European mole)4
Sorex araneus (common shrew)3
Sorex minutus (Eurasian pygmy shrew)1
Sorex minutissimus (Eurasian least shrew)2
Crocidura russula (greater white-toothed shrew)3
Oryctolagus cuniculus 32.5 kg
Lepus europaeus (European hare)43.4 kg
Lepus timidus (mountain hare)2
Ochotona pusilla (steppe pika)3
Myotis myotis 125.7 g
Pipistrellus pipistrellus 14.4 g
Bufo bufo 2
Bufo calamita 4
Rana dalmatina 1
Rana ridibunda 1
Rana temporaria 1
Lacerta viridis 1
Anguis fragilis 2
Coronella austriaca 2
Malpolon monspessulanus 1
Natrix maura 1
Vipera berus 1
Cervus elaphus (red deer)15
Capreolus capreolus (roe deer)3
Bison priscus (bison)56
compared to subspecies mediator
Equus cf. mosbachensis (horse)11
Hemitragus cedrensis (tahr)10
Palaeoloxodon antiquus (elephant)1
Rupicapra sp. 1
Canis lupus lupus (gray wolf)143.0 kg
Vulpes vulpes (red fox)2
Felis silvestris (European wildcat)14.5 kg
Ursus arctos (brown bear)1
Mustela nivalis (least weasel)1061.7 g